YosHINo , 1975 ( PeRCIfoRMes : lABRIdAe ) , fRoM NoRTH sulAWesI , INdoNesIA

One specimen of Bodianus izuensis Araga and Yoshino, 1975, and three specimens of B. masudai Araga and Yoshino, 1975, were obtained from fish markets in Bitung and Kema, Minahasa, North Sulawesi, Indonesia, between March–July 2009. The specimens were caught from depths of about 20–30 m at coral reef ecosystem around Lembeh Island and Kema. This is the first record of these species in Indonesian waters. Morphological features, diagnostic characters, and distribution are discussed and illustrated, and color photographs of the species are presented.


INTRoduCTIoN
Labridae commonly known as wrasse consist of about 57 genera and 575 species (Parenti and Randall, 2000) is one of the most dominant fish families in Indonesia and its adjacent waters.Wrasses are small to medium-sized marine fishes, mostly tropical, associated with coral reefs; some species have a warm temperate distribution.One of the genera in the Labridae is Bodianus, characterized by the possession of a prominent frontal shelf on the neurocranium, posteriorly roofing a distinct medial ethmoid-frontal depression, and a sharply angled anteroventral profile of the dentary with an interdigitating joint at the symphysis (Gomon, 2006).Although most species of Bodianus are found in relatively shallow waters a few live in greater depths (Gomon, 2006), such as Bodianus sp.recorded by ROV Fukushima Aquarium in more than 150 m at Manado, North Sulawesi (Iwata, pers. comm.) and Bodianus masudai collected from rocky reefs at 30-113 m depth (Gomon, 2006).
Most labrids are protogynous hermaphrodites and have three distinct colour patterns correspond-ing with age and sexual development.Gomon (2006) described the ancestral color pattern of Bodianus as follows: "juveniles darkly pigmented with 5 or 6 narrow pale bands on the body posterior to the head; a prominent black spot anteriorly and posteriorly on dorsal fin, a third somewhat centrally on the fleshy caudal-fin base, a fourth posteriorly on the anal fin, a fifth on the fleshy pectoral-fin base and a sixth on the pelvic fin; initial-phase adults with red, horizontal stripes; initial-and terminal-phase adults with vermiculations around the eye".
The first specimen of Bodianus Bloch, 1790 (type species Bodianus bodianus) species is collected from Fish Market, Jakarta, by Hardenberg and Hubbs on 6 May 1929 and now is deposited at University of Michigan Museum of Zoology (UMMZ), Ann Arbor, Michigan, USA (Eschmeyer, 2012).The genus Bodianus, one of the genera in the family Labridae, comprises 43 species representing 10 subgenera, with the greatest diversity in the Indo-West Pacific (Gomon, 2006).
Ten species of fish of genus Bodianus (B. anthioides, B. axillaris, B. bilunulatus, B. bimaculatus, B. frenchii, B. leucosticticus, B. loxozonus, B. mesothorax, B. perditio and B. sanguineus) have been reported from Indonesia (Eschmeyer, 2012).Additional two species of B. izuensis and B. masudai from Kema, North Sulawesi, reported herein are bringing the total number of species of this genus known in Indonesian waters to 12.This number of species is relatively high compared to those reported from several other localities, so far only eight species were reported from South China Sea (Randall and Lim, 2000) and four species from southwestern Thailand, Andaman Sea (Satapomin, 2011).

MATeRIAls ANd MeTHods
Four specimens of pigfish were obtained from Girian Fish Market, Bitung, North Sulawesi.The specimens were immediately photographed, labeled and preserved in formalin solution 10% and deposited at the Technical Implementation Unit for Marine Biota Conservation, Indonesian Institute of Sciences, Bitung, for further study.Valid record of the fish specimen referred to the data from Eschmeyer (2012).
Counts and measurements generally followed those of Hubs and Lagler (1947) with some modifications (Gomon, 2006).Standard length and total length were abbreviated as SL and TL, respectively.Measurements were made on the left side of specimen using dial-point caliper to the nearest of 0.01 mm.Proportional measurements on body and head are presented as percentages of standard length (SL) and head length (HL), consecutively.
Remark -Bodianus izuensis resembles B. bimaculatus in having a less pointed snout and a smaller maximum size (Masuda et al., 1975) and differs from B. bimaculatus by having prominent black stripes on the body.Based on the size and color of the specimens examined, these two species from Bitung and Kema, North Sulawesi are categorized to terminal-phase adult.No accurate data on handline depth of this species, however judging the method used by fishermen collecting the specimen, B. izuensis possibly were collected from shallow waters rocky reef.The same species has been reported from off Shizuoka Prefecture, Japan was collected at depths of about 30-35 m on deep rocky reefs (Masuda et al., 1975).

Bodianus masudai Araga et Yoshino, 1975,
Pig fish (Fig. 2; Tabel 2) Bodianus masudai Araga andYoshino, 1975 in Masuda et al., (1975): 297, pl. 103-H;Masuda et al. (1984): 203, pl. 195-O;Shao, 1986: p. 182  distribution -Bodianus masudai, initially known only in Japan, Norfolk Island, Taiwan and New Caledonia (Masuda et al., 1975;Shao, 1986;Gomon, 2006;Fourmanoir in Gomon, 2006) can be considered as species with an antitropical distribution.Thus, the present specimen from Bitung, Northern Sulawesi, Indonesia, represents the first record of this species from the tropical region in the Western Pacific.It suggests that the species is widely and continuously distributed to the western Pacific from Japan, i.e. from Indonesia to Southern Hemisphere.The Indonesian specimen was collected at 20-30 m depth, whereas in Japan, New Caledonia and Norfolk Island this species were collected from 30-113 m and may be more common at greater depths (Gomon, 2006).Further collection of the specimens from Indonesia are required to determine whether the Indonesian population also occurs in deeper water.
Remark -Bodianus masudai resembles to B. neopercularis, B. opercularis and B. sepiacaudus in having a prominent red striped color pattern in adults, but differs from them in having the opercular spot reaches below the upper end of the pectoral-fin base and in having fewer lateral-line scales.B. masudai also differs from these three species in having the lowermost red body stripe of adults originating on the ventral edge of the eye rather than on the underside of the jaw.Following Gomon (2006), all specimens of B. masudai collected from Bitung and Kema, North Sulawesi can be categorized to terminal-phase adult.